Crossword clues for ___ on the knuckles 4 letters:FEARCause of white knucklesSLUGGive a knuckle sandwichSLAPPlay with the knuckle, as strings on a bass guitarBARE___-knuckle (not wearing boxing gloves)FISTKnuckle sandwich ingredient?SLUGGive a knuckle sandwich toFISTBare-knuckle boxing weaponSEGASonic & Knuckles game companyRAPSKnocks with one's knucklesPIGSKnuckles or feetHEADWord with war or knuckleBAREWord with back or knuckleAPESMany knuckle-walking creaturesCAVEKnuckle underHANDKnuckle's placeBAREPrecedes foot and knuckleSEGASonic & Knuckles publisherFISTKnuckle sandwichBARELead-in to bones or knucklesSEGAKnuckles the Echidna's companySLUGPresent with a knuckle sandwichBAREWord before "bones" or "knuckle"ARAPGet __ on the knucklesOBEYKnuckle underBARELead-in to "bones" or "knuckle"ARAP___ on the knucklesHOCKHam ___ (pork knuckle)BOORKnuckle-draggerBAREWord before bones or knucklesLOUTKnuckle-draggerRAPSKnuckle knocksARAP__ on the knucklesGIVEKnuckle underRAPSUses the knucklesHOYTKnuckle ball legend WilhelmBOWSKnuckles downBARE___-knuckle boxingAPESKnuckle draggersKNUCKLEPart of a handKNUCKLEFinger jointKNUCKLECertain jointKNUCKLE___ under (capitulate)KNUCKLESFist partsKNUCKLEWork hard ("down")KNUCKLEPart of a fistKNUCKLEKind of ball or sandwichKNUCKLEType of pitched ball.KNUCKLEGet serious, with "down"KNUCKLEDGot to work, with "down"KNUCKLED__ under (gave in)